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Follow-up studies combining mitochondrial metabolite profiling, organelle transport assay and structure-guided mutagenesis identified a critical role of SLC25A39 in mitochondrial glutathione uptake, a discovery that has also been independently reached by a recent report 14

Previous work on Hld Sa has shown that it self-aggregates and was originally purified as a 270 kDa protein complex, consistent with observation of the 3 kDa peptide having an activity present in the greater than 100 kDa fraction